Journal Articles & Other Academic Publications

Check-in often! We update this page as soon as an IPREFER collaborator provides us with new information. All authors in bold text are affiliated with the IPREFER project.


  • Zhang, Julia, Greatens, Nicholas, Arenz, Brett, Flynn, Jennifer, & Anderson, James. (2024). First report of powdery mildew caused by Erysiphe cruciferarum on field pennycress (Thalaspi-arvense) in North America. Plant Disease. [Manuscript submitted for publication on May 5, 2024].
  • Rocha, Leonardo. “Improving the Management of the Soybean Cyst Nematode (*Heterodera Glycines* Ichinohe): From Field Practices to Next-generation Sequencing Approaches.” Doctoral, Southern Illinois University, 2024.
  • Forcella, Frank, M. Petersen, William L. Perry, Samantha S. Wells, Alexander Hard, Russ W. Gesch, Yesuf A. Mohammed, Cody Hoerning, Tad L. Wesley, E. Ambrosi & Winthrop B. Phippen. 2024. Flies associated with floral canopies of the new oilseed crop, pennycress, in the Midwestern U.S.A. The Great Lakes Entomologist 56: 154-165. Open Access)
  • Gomes, Victor Emmanuel de VasconcelosAnn M. Kennedy, Amanda R. Darcy & Alexander J.  Lindsey. Hormone type, soaking duration and seed coating affect dormancy and germination of pennycress (Thlaspi arvense) lines. Seed Science and Technology. (Open Access).
  • Perez, Jayleen, Cubins, Julijia. A., Samantha S. Wells, Gregg A. Johnson, Katie L. Black, Ariana Gonch, Frank Forcella, & Russ W. Gesch. 2024. Stover removal has minimal impact on pennycress seeded into standing corn. Crop Science, 1–9,
  • Yang, Liangcheng, Tuba Yasmin Lubna, Michael A. Moklak, Barsanti Gautam, Nicholas J. Heller, Robert L. Rhykerd, David E. Kopsell & John C. Sedbrook. (2024) Impacts of harvest date and concurrent alkali pretreatment and ensiling on anaerobic digestion of pennycress biomass. Fermentation 10: 96.


  • Al Sulaimi, Rawan, Ali Macknojia, Mohammad Eskandari, Asghaar Shirani, Barsanti Gautam, Wonkeun Park, Payton Whitehead, Ana Paula Alonso, John C. Sedbrook, Kent D. Chapman, Diana Berman, 2023. Evaluating the Effects of Very Long Chain and Hydroxy Fatty Acid Content on Tribological Performance and Thermal Oxidation Behavior of Plant-Based Lubricants. Tribology International, p.108576.
  • Forcella, Frank, Zachary M. Portman, Samantha S. Wells, William Perry, Russ Gesch, Cody Hoerning, Tad Wesley & Winthrop B. Phippen. (2023). Abundance and diversity of bees visiting flowering pennycress, a new oilseed crop in the midwestern USA. The Great Lakes Entomologist 56 (1):99-107.
  • Koirala, Nasib, David J. Barker, Russ W. Gesch, Nicholas J. Heller, Alexander W. Hard, Samantha S. Wells, Winthrop B. Phippen, Alexander J. Lindsey. 2023. Seed pelleting and storage effects on germination of Pennycress (Thlaspi arvense L.) Crop Sci. First published: 10 August 2023. OPEN ACCESS
  • Meyer, Ryan T., Nicholas J. Heller, Robert L. Rhykerd, Alex W. Hafner & William L. Perry. 2023. Biomass, nitrogen, and carbon loss from the novel oilseed pennycress relative to annual ryegrass and cereal rye. Cogent Food & Agriculture, 9 (1) (Article Published online: 24 May 2023
  • Mohammed, Yesuf, Russ Gesch, Samantha Wells, Nicholas Heller, Alex Lindsey, Alexander Hard, Winthrop Phippen & Bethany Bedeker. 2023. Yield tradeoffs of early corn harvest to enhance pennycress establishment. Agron. J. 115:4 (May 5, 2023) 1762-1775. Doi: 10.1002/agj2.21370


  • Koirala, Nasib, David Barker, Carin A. Helfer, Winthrop B. Phippen, Nicholas Heller, Alexander W. Hard, Wells, Samantha & Alexander J. Lindsey. A process to enhance germination of a wild pennycress variety. (2022). Seed Science & Technology, Vol. 50 (2), August 2022, 195-205(11). OPEN ACCESS
  • Nunn, Adam, Isaac Rodríguez-Arévalo, Zenith Tandukar, Katherine Frels, Adrián Contreras-Garrido, Pablo Carbonell-Bejerano, Panpan Zhang, Daniela Ramos-Cruz, Katharina Jandrasits, Christa Lanz, Anthony Brusa, Marie Mirouze, Kevin Dorn, Brice Jarvis, John Sedbrook, Donald L. Wyse, Christian Otto, David Langenberger, Peter F. Stadler, Detlef Weigel, M. David Marks, James A. Anderson, Claude Becker & Ratan Chopra (corresponding author). 2022. Chromosome-level Thlaspi arvense genome provides new tools for translational research and for a newly domesticated cash cover crop of the cooler climates. Plant Biotechnology. doi: OPEN ACCESS
  • Phippen, Winthrop B., Rob Rhykerd, John C. Sedbrook & Christine Handel. 2022. From farm to flight: CoverCress as a low carbon intensity cash crop for sustainable aviation fuel production. A review of progress towards commercialization. Front. Energy Res., 24 June 2022. Sec. Bioenergy and Biofuels. 10: 2022,
  • Tandukar, Zenith, Ratan Chopra, Katherine Frels, Brett Heim, M.David Marks & James A. Anderson. 2022. Genetic dissection of seed characteristics in field pennycress via genome-wide association mapping studies. The Plant Genome, 00, e20211.
  • Verhoff, Kyle, Win Phippen, Nicholas Heller, Alex Lindsey. 2022. Winter-type oilseed pennycress crop staging guide. Crop, Forage, and Turfgrass, 8, e20165. See the accompanying Winter-Type Oilseed Pennycress Staging Guide.
  • Yang, Liangcheng, Lucas D. Lamont, John C. Sedbrook, Nicholas J. Heller and David E. Kopsell. (2022) Anaerobic Digestion of Cereal Rye Cover Crop. Fermentation 8(11):617.


  • Betts, K., Wells, Samantha S., Sarangi, D., Bernards, Mark L., Wyse, D.L. 2021. Field pennycress (Thlaspi arvense) response to herbicides applied postemergence. Weed Science Society of America, Annual Meeting Proceedings 60:20.
  • Esfahanian, Maliheh, Tara J. Nazarenus, Meghan M. Freund, Gary Mcintosh, Winthrop B. Phippen, Mary E. Phippen, Timothy P. Durrett, Edgar B. Cahoon & John C. Sedbrook. 2021. “Generating pennycress (Thlaspi arvense) seed triacylglycerols and acetyl-triacylglycerols containing medium-chain fatty acids.” Energy Res. January 28, 2021. doi: 10.3389/fenrg.2021.620118. The Supplementary Material for this article can be found online at:
  • Hagaman, Matthew, Darner, Rebekka, Sparks, Rachel, Schoenfelder, Emily & Wm. Hunter. “Cover Crop Science Project Book: An Inquiry-based Project Book for 4-H Youth.”
  • Jarvis, Brice A., T.B. Romsdahl, Michaela McGinn, T.J. Nazarenus, E.B. Cahoon, K.D. Chapman & John C. Sedbrook. (2021). “CRISPR/Cas9-induced fad2 and rod1 mutations stacked with fae1 confer high oleic acid seed oil in pennycress (Thlaspi arvense L.).” Frontiers Plant Sci. 12:652319.
  • Marks, M.David, Ratan Chopra, & John.C. Sedbrook. (2021). “Technologies enabling rapid crop improvements for sustainable agriculture: Example pennycress (Thlaspi arvense L.).” Emerg. Topics Life Sci. ETLS20200330.
  • Ott Matthew A., G. Gardner, Krishna M. Rai, D.L. Wyse, M.David Marks & Ratan Chopra. (2021). TRANSPARENT TESTA 2 allele confers major reduction in pennycress (Thlaspi arvense L.) seed dormancy. Industrial Crops & Products. 174 (2021) 114216. OPEN ACCESS:


  • Chopra, Ratan, Folstad, Nicole & M. David Marks. (2020). “Combined genotype and fatty-acid analysis of single small field pennycress (Thlaspi arvense) seeds increases the throughput for functional genomics and mutant line selection.” Ind Crops Prod., 156: 112823.
  • Chopra, Ratan, Evan. B. Johnson, Ryan Emenecker, Edgar B. Cahoon, Joe Lyons, Daniel J. Kliebenstein, Erin Daniels, Kevin M. Dorn, Maliheh Esfahanian, Nicole Folstad, Katherine Frels, Michaela McGinn, Matthew Ott, Cynthia Gallaher, Kayla Altendorf, Alexandra Berroyer, Baraem Ismail, James A. Anderson, Donald L. Wyse, Tim Ulmasov, John C. Sedbrook, & M.David Marks. (2020). “Identification and stacking of crucial traits required for the domestication of pennycress.Nat Food, 1 (2020): 84–91. This paper was the subject of a “News & Views” article: Britt, Anne B. “From stinkweed to oilseed.” Nat Food 1: 84-91.
  • Sedbrook, John, C. & Timothy P. Durrett. (2020). “Pennycress, carbon wise: Labeling experiments reveal how pennycress seeds efficiently incorporate carbon into biomass.” J. Exp. Botany, 71: 2842-2846.
  • Tyl, Catrin, Lee DeHann, Katherine Frels, Prabin Bajgain, M.David Marks & James A. Anderson. (2020). “Emerging crops with enhanced ecosystem services: Progress in breeding and processing for food use.” Cereal Foods World, 65, 2 DOI:


  • Cubins, Julija, A., Samantha S. Wells, Katherine Frels, Matthew A., Ott, Frank Forcella, Gregg A. Johnson, Mainder K. Walia, Roger L. Becker, & Russ W. Gesch. (2019). “Management of pennycress as a winter annual cash cover crop: A review.Agron. Sustain. Dev. 39, 46 (2019): 46, OPEN ACCESS.